Perona
Perona is a ghost princess who serves as one of the commanders under Gecko Moria on Thriller Bark. She directs zombie forces and uses her abilities to weaken intruders while maintaining a network of ghosts for surveillance. Her allegiance lies with Moria and his crew, and her goals center on protecting the island's operations and later seeking a preferred dark environment after events force relocation. She is known for her command over Negative Hollows and her distinctive floating form during confrontations.
Throughout her time on Thriller Bark she coordinates with Absalom and Hogback while overseeing subordinates like Kumashi. After forced departure she ends up on Kuraigana Island where she encounters Zoro and later interacts with Mihawk. Her later activities include monitoring news about Moria and deciding on further travel.
Appearance
Perona appears in a frilly dress paired with striped leggings and a crown-like headpiece that accentuates her pigtails. She maintains this style across multiple locations including the halls of Thriller Bark and later settings on Kuraigana Island. Her posture often involves floating with arms at her sides during encounters.12
Additional details include a large bow and long light hair that frame her face during moments of shock or determination. She carries a cross-shaped staff in certain scenes and holds small objects while surrounded by her ghost companions.34
Personality
Perona expresses a strong preference for cute subordinates only and reacts with irritation when expectations are unmet. She maintains authority through fear of punishment among the zombies under her command and shows playful yet eerie laughter during confrontations.56
She displays shock and denial when plans fail or unexpected figures appear and later voices frustration over the absence of desired comforts in new surroundings. Her tone shifts between commanding and panicked depending on the situation.78
Abilities and powers
She demonstrates the ability to float freely and pass through walls as part of her ghost-like powers. Her body can separate into a floating head and ghost form while physical strikes pass harmlessly through her during combat.910
Perona activates techniques such as Ghost Wrap and Special Hollow while launching explosive attacks from midair. She maintains advantage through intangible ghost assaults and recognizes true body locations during engagements.1112
History
Thriller Bark operations
On Thriller Bark Perona directs the Straw Hats toward a prepared bath and later discusses their bounties with Absalom and Hogback. She arrives carried by Kumashi near the clock tower and leads wild and surprise zombies at midnight while maintaining a ghost network for reporting enemy movements.135
On Thriller Bark Perona stands beside an ornate bath entrance and invites a dirty group inside while gesturing toward the ready facility. She later sits on a throne in a large hall while directing zombie subordinates and receives reports about captured intruders from her network.141516
Perona finds Absalom hiding inside her room after deciding to take food supplies from an enemy ship and hides inside a protective bubble with Kumashi and Humming when an issue arises. She regains consciousness in a ruined hall after the enemy departs and witnesses Oars fist impact before choosing to flee on the pirates ship.1718
Kuraigana Island period
Relationships
Perona maintains close ties with Kumashi who accompanies her during operations and attempts to report issues while she hides with allies inside a protective bubble. She also interacts with Absalom who hides in her room and with Hogback during discussions of enemy bounties.1721
Perona watches enviously as cute bear cubs like Mihawk and later argues with the swordsman over her contributions to farm work before deciding to depart. She excitedly shares news of Moria's survival with Mihawk while crying out about what happened to her master.2223
Major battles
Perona engages Usopp in a throne room corridor where she launches Negative Hollows and Rokuren Mamushi Boshi techniques. She phases through physical contact and activates Ghost Wrap before being struck by an explosive star attack that forces her to recognize the trap.1112
Perona confronts Bartholomew Kuma at the Thriller Bark castle gates and uses Negative Ghost and Special Hollow attacks against him. She suddenly vanishes from the deck after questioning his identity and connection to Moria.248
Thriller Bark
Perona launches Rokuren Mamushi Boshi from midair against Usopp and activates Ghost Wrap after summoning Mini Hollow ghosts while grappling and phasing through the opponent. She later realizes her attacks succeeded because the target fled rather than phasing through walls.2526
Perona is attacked by cockroaches at the base of the Tower of Justice and faces Franky who wields a massive hammer before being defeated inside the collapsing hall. She regains consciousness later amid the ruined surroundings.2728